1. Introduction: From Mechanism to Vitalism and Back; 2. Useful Causes: The Quest for Inducers and Organizers; 3. Idealized Causal Concepts as Research Tools: Morphogens; 4. Causes in Harmony; 5. Epilogue: Understanding, Progress, and Manipulability; References.

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This Element examines causality, mechanisms, explanations and discovery in classical experimental embryology as well as molecular developmental biology.

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